Journal Article stripe rust Aegilops speltoides Research Support, Non-U.S. Gov't Thinopyrum scirpeum asymmetric spindle candidate gene centromere chromosomal substitution chromosome drive chromosome nondisjunction chromosome translocation line flow cytometry genome-wide association study genotyping by sequencing molecular marker pollen grain mitosis powdery mildew quantitative trait loci supernumerary B chromosome tetraploid Thinopyrum elongatum wheat yield-related