Journal Article species distribution models Research Support, Non-U.S. Gov't climate change Centaurea stoebe (spotted knapweed) Himalayas Quaternary climatic changes Ramsar Soil adaptation animals biodiversity conservation biotic interactions biotic modifiers climate change scenarios climate suitability climatic niche colonization ability conservation cryptic invasion demography ecosystems facilitation future projections herbarium specimens isolation by distance light competition microorganisms niche model niche truncation