Journal Article Research Support, Non-U.S. Gov't administración ambiental benthic biodiversity environmental stewardship fully protected areas irradiance macroalgae marine ecology marine protected areas mixed methods métodos mixtos partially protected areas polar regime shift sistemas socioecológicos social-ecological systems áreas completamente protegidas áreas marinas protegidas áreas parcialmente protegidas 完全保护区 海洋保护区 混合方法 环境管理 社会生态系统 部分保护的保护区