Journal Article Research Support, Non-U.S. Gov't Christmas Island red crab El Niño-Southern Oscillation Gecarcoidea natalis Research Support, U.S. Gov't, Non-P.H.S. Southern Oscillation Index climate change fragmentación fragmentation graph theory grasslands land crab modelos de redes network models pastizales phenology population size redes ponderadas tamaño poblacional teoría de gráficos weighted networks 加权网络 图论 破碎化 种群大小 网络模型 草地