African forest elephant Gorilla gorilla gorilla Journal Article Loxodonta cyclotis Pan troglodytes troglodytes Research Support, Non-U.S. Gov't Research Support, U.S. Gov't, Non-P.H.S. Syncerus caffer nanus betweenness centrality central chimpanzee centralidad intermedia forest buffalo modelo de ocupación occupancy modeling western lowland gorilla 中部黑猩猩 中间中心性 占有建模 森林野水牛 西部低地大猩猩 非洲森林象