Journal Article Mitogen-Activated Protein Kinases 6M3C89ZY6R Alkaloid biosynthesis Cyclopentanes EC 2.7.11.24 EC 3.1.3.16 MAP kinase Nicotine Oxylipins Phosphoprotein Phosphatases Plant Proteins Research Support, Non-U.S. Gov't Transcription Factors gene regulation hairy roots nicotine phytohormones post-translational regulation protein phosphatase 2C secondary metabolism specialized metabolites sucrose non-fermenting-1 (SNF)-related protein kinases tobacco transcription factors