Journal Article Research Support, Non-U.S. Gov't Rhopalocera Soil allozyme electrophoresis atmospheric nitrogen butterflies climate change climatic oscillations community structure comportamiento de dispersión dispersal behavior early spring ecological tolerance estructura comunitaria generations genetic structure glacial refugia global change global warming long-term observations moths mountain archipelagos nitrógeno atmosférico postglacial colonization procesos estocásticos prolonged vegetation period remplazo temporal de especies serie de tiempo species distribution models