Journal Article Alaska Research Support, U.S. Gov't, Non-P.H.S. behavioral plasticity biotelemetry biotelemetría caribou climate change collective memory comportamiento multidimensional ecological risk management especie migratoria gestión de riesgos ecológicos hidden Markov models leatherback turtle migratory range migratory species modelos ocultos de Markov multidimensional behavior snow survivorship temperature tortuga laúd wind 多维行为 棱皮龟 洄游物种 生态风险管理 生物遥测 隐马尔可夫模型