Journal Article N‐mixture model N‐混合模型 Research Support, Non-U.S. Gov't agro-ecology behavioral variation bird cambio de uso de suelo climate change ecología tropical environmental gradient homogenization land‐use change modelo N‐mixture multiple stressors protected area tropical tropical ecology turnover variación conductual área protegida 保护地 土地利用变化 热带生态学 行为变异 补偿